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Veterinary ophthalmologists are often asked, "How well do
animals see?" Visual function involves a combination of many factors,
including: the field of view, depth perception (ability to judge distances),
acuity (focusing ability), perception of motion, and color differentiation.
All of these functions must then be integrated by the brain to produce
useful vision. Although we are unable to ask our pets to read an eye chart,
through comparative studies, it is possible to make some educated
assumptions about their vision. |
The position of the eyes within the head determines the
degree of peripheral vision as well as the amount of the visual field that
is seen simultaneously with both eyes. This binocular vision is necessary
for judgment of distances. Dogs have eyes which are placed on the sides of
the head, resulting in a visual field of 240 degrees compared with the human
field of 200 degrees. The central, binocular field of vision in dogs and
cats is approximately half that possessed by humans.
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The
eyes of dogs and cats have many of the modifications typically seen in
animals which evolved as nighttime hunters. The pupil functions much as the
aperture for a camera and can dilate for maximal light capturing ability in
dogs and cats. In addition, there is a reflective layer under the retina
called the tapetum which serves to intensify vision in dim light. The
"mirror" effect of the tapetum results in the "eye shine" observed when an
animal looks into a car's headlights. While dim light vision is enhanced by
the tapetum, scattering of the reflected light may result in reduced acuity.
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| Although it is commonly believed that dogs and cats see only
in black and white, recent evidence suggests that animals may have some
degree of useful color vision. The perception of color is determined by the
presence of cone photoreceptors within the retina. These cone cells function
in bright light conditions and comprise approximately 20% of the
photoreceptors in the central retina of the dog. In humans, the central
retina (macula) is 100% cones. Behavioral tests in dogs suggest that they
can distinguish red and blue colors but often confuse green and red. |
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Human vision with full color range possible because of specialized retina including cone photoreceptors and macula.
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Dogs and cats have vision with a very limited color spectrum. Blue is visible but red, green, and yellow all appear similar.
Animals rely on contrast and movement to identify objects. This type of normal vision is restored following cataract surgery with implantation of a replacement lens.
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Vision through a cloudy, cataractous lens.
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Following removal of a cataract without replacement lens implantation,
vision is improved but still blurry.
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Acuity is the ability to focus so that two objects appear as
distinct entities. This is the value that is measured in people using an eye
chart. In animals (and in infants) this can be measured using retinoscopy.
Visual acuity in normal dogs has been estimated to be 20-40% that of humans.
This means that, at 20 feet, a dog can distinguish an object that a human
could see at 90 feet. Acuity is a function of the clarity of the structures
of the eye (cornea, aqueous humor, lens, and vitreous) as well as the
combined refractive powers to focus the image clearly on the retina. One of
the most common diseases which can affect the clarity of the eye is
formation of a cataract within the lens. Cataracts in dogs are most often
inherited and
may
affect dogs at any age. As the lens becomes progressively opaque, an
animal's vision deteriorates so that only light and dark perception exists.
Surprisingly, animals function relatively well in familiar surroundings,
even with severe vision impairment. This illustrates the ability of dogs and
cats to depend heavily on their other senses, namely smell and hearing. |